Dr. Gregory Lewis' decision to pursue a career in ecology was influenced strongly by his early work as an undergraduate research assistant at the University of Georgia's Savannah River Ecology Laboratory (SREL). At SREL, he assisted in research by Dr. Barbara Taylor on zooplankton in artificial reservoirs and in Carolina bay wetland ponds, as well as in studies by Dr. J. Vaun McArthur of stream bivalves and leaf decomposition. Lewis also conducted research with Dr. Wade Worthen on the influence of ant predation on the survival of Drosophila larvae developing in mushrooms. After earning his bachelor of science in biology from Furman, he began his doctoral studies in ecology and evolutionary biology at Cornell University, where he was supported in part by a National Science Foundation Graduate Fellowship. His doctoral research, under the supervision of Dr. Gene Likens (of the Cary Instit ute of Ecosystem Studies) and Dr. Bob Howarth, examined the influences of forest insect defoliation, forest species composition, and soil types on stream chemistry in the Allegheny National Forest of northwestern Pennsylvania. After completing his Ph.D., Lewis worked as a postdoctoral research associate with Dr. Barbara Taylor at SREL, examining the long-term fate of radiocesium in artificial reservoirs that had received heated effluents from nuclear reactors. Dr. Lewis joined the Furman Biology faculty in 2000.

In addition to teaching and research, Dr. Lewis has led or participated in several study away programs in the Biology Department, including travel to Costa Rica (Tropical Ecology), Ecuador (Biology of the Andes and Galápagos Islands), and Belize (Marine Biology). He also is an Affiliate Faculty member with the David Shi Center for Sustainability at Furman.

Name Title Description


Seminar in Biology

Presentations of current topics in biology by students, faculty, and visiting scientists. Emphasis is on effective oral communication and critical examination of scientific information and ideas.


Biology and the Environment

The basic principles common to living organisms, with emphasis on environmental biology. Designed for non-science majors. Topics include: ecology, evolution, biodiversity, and environmental issues, plus a synopsis of cellular biology, genetics, and physiology.


Foundations of Biology

Introduction to the unifying concepts of biology. Topics include: cell structure and function, metabolism, genetics, evolution, and the diversity of life. Laboratories emphasize an investigative approach. Designed for science majors.


Research and Analysis

Introduction to purposes and methods of scientific inquiry. Topics include: philosophy of science, research design, use of biological literature sources, fundamental laboratory techniques, statistical analysis, and survey of careers in biology. Laboratory includes designing, performing, and reporting on research projects.



The interactions between organisms and their environments, and the consequences of these interactions for population dynamics, community structure, and the flow of energy and matter through ecosystems. Also, consideration of environmental issues and conservation. Laboratories include local field work, experiments, and possibly one or two overnight/weekend field trips.


Environmental Systems

Interdisciplinary examination of the physical, biological and chemical processes that control the flow of matter and energy in surface environments on planet Earth. Emphasis on the interactions between abiotic and biotic processes. Lab includes field studies and weekend field trips.


Infectious Disease Ecology

The impacts of ecosystem changes on emerging and reemerging infectious diseases in animals and plants (both domestic and wild) and how these diseases influence communities, ecosystems, and human welfare. Laboratories include field and lab work.


Tch Mthds & Mtrls in Biology

Practicum for developing teaching skills in biology, including the laboratory and field work. Topics include planning and preparation, safety and storage, and instruction. Practicum will provide experiences teaching in the university classroom.


Fly Fishing and Rivers

Introduction to Fly Fishing. Watershed geology, stream structure, reading a stream to identify likely location of fish, life cycle of aquatic insects and other food sources, biology of fish and common fish behavior, resource conservation, ethics of catch and release"


Environment and Society

Interdisciplinary examination of the causes, potential solutions and ethical dilemmas associated with environmental problems on various spatial, temporal, political and social scales (individual to global).


Pristine Nature: Myth Or Real

An examination of the concepts of wilderness and pristine" nature from scientific

Dr. Lewis conducts research within the River Basins Research Initiative, which involves close collaboration with students and fellow faculty in the Biology and Earth and Environmental Science Departments. Broadly speaking, Dr. Lewis is interested in how land cover and human activity within watersheds influences stream and river ecosystems. He is especially interested in how land cover (both urban and rural) influences the biogeochemistry of elements such as nitrogen and carbon in watersheds in the piedmont and Blue Ridge regions of South Carolina. Although he focuses on patterns in stream chemistry, he also is interested in how air pollution (in both rain and dust) may enhance the supply of nitrogen and other elements to streams in urban watersheds. Dr. Lewis also collaborates with Dr. Min-Ken Liao on studies of how both point and non-point sources of pollution in urban areas affect the abundance and genetic diversity of antibiotic-resistant bacteria in headwater streams. Further, Dr. Lewis has conducted research on the influence of wastewater treatment plant effluent on river chemistry and on riparian vegetation and woody debris in urban and rural streams.

* Denotes undergraduate coauthor

  • Lewis, G. P., C. B. Andersen, T. P. Moloney*, and S. Muthukrishnan. 2015. Relationships between dry deposition of ions and urban land cover in the South Carolina piedmont. Water, Air, and Soil Pollution 226:266
  • Anderson, B. W.*, S. McCauley*, G. P. Lewis, and M.-K. Liao. 2014. Impacts of a poultry processing plant on the diversity of Escherichia coli populations and transferability of tetracycline resistance genes in an urban stream in South Carolina. Water, Air and Soil Pollution 225:2030.
  • Andersen, C.B., G. P. Lewis, M. Hart*, and J. Pugh*. 2014. The impact of wastewater treatment effluent on the biogeochemistry of the Enoree River, South Carolina, during drought conditions. Water, Air and Soil Pollution 225:1955.
  • Dripps, W., G. P. Lewis, R. Baxter*, and C. B. Andersen. 2013 Hydrogeochemical characterization of headwater seepages inhabited by the endangered Bunched Arrowhead (Sagittaria fasciculata) in the upper piedmont of South Carolina. Southeastern Naturalist 12(3)619-637.
  • Williams, A.*, C. B. Andersen, and G. P. Lewis. 2009. Evaluating the effects of sample processing treatments on alkalinity measurements. Journal of Hydrology 377:455-464.
  • Muthukrishnan, S., G. P. Lewis, and C. B. Andersen. 2007. Relations among land cover, vegetation index, and nitrate concentrations in streams of the Enoree River Basin, piedmont region of South Carolina, USA. In D. Sarkar, R. Datta, and R. Hannigan, editors. Developments in Environmental Science, vol. 5. Elsevier.
  • Lewis, G. P., J. Mitchell*, C. B. Andersen, D. C. Haney, M.-K. Liao, and K. A. Sargent. 2007. Influence of urbanization on stream chemistry and biology in the Big Brushy Creek watershed, South Carolina, USA. Water, Air and Soil Pollution 182:303-323.
  • Lewis, G. P., and G. E. Likens. 2007. Changes in stream chemistry associated with insect defoliation in a Pennsylvania hemlock-hardwoods forest. Forest Ecology and Management 238:199-211.
  • Andersen, C.B., G. P. Lewis, and K. A. Sargent. 2004. Influence of wastewater treatment effluent on concentrations and fluxes of solutes in the Bush River, South Carolina, during extreme drought conditions. Environmental Geosciences 11:28-41.
  • Lewis, G. P., and G. E. Likens. 2000. Low stream nitrate concentrations associated with oak forests on the Allegheny High Plateau of Pennsylvania. Water Resources Research 36:3091-3094.
  • Lewis, G. P., B. E. Taylor, J. E. Pinder III, and P. M. Dixon. 2000. Apparent decline of the sediment 137Cs inventory of an abandoned reactor cooling reservoir: export or uncertainty? Journal of Environmental Radioactivity 49:293-306.
  • Lewis, G. P., and W. B. Worthen. 1992. Effects of ant predation and mushroom desiccation on survivorship of mycophagous Drosophila tripunctata larvae. Oikos 64:553-559.
Cornell University
Furman University

Connect with Admission

Furman is one of the nation's premier liberal arts and sciences universities. We offer our students The Furman Advantage—an over-arching approach to education that promises every student a four-year personalized pathway, a team of advisors and mentors, and the opportunity for an engaged learning experience that is tracked and integrated with the students' academic and professional goals.

Want more information about the admission process at Furman?

Contact us

Once you see our campus, making the right college decision will be so much easier.

Plan a visit

Undergraduate Evening Studies provides adults the opportunity to receive an education from one of the premier liberal arts universities in the nation.

Whether you are starting or continuing your education, or have been away from the classroom for a few months or several years, our program provides many services to assist you with accomplishing your educational and professional goals.

Apply now

Our graduate studies program is designed for the professional educator.

We know the challenges teachers and administrators face every day, and we are committed to helping you become a leader within your school system or district.

Apply now
  • Furman University